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1. D.V. VAYENAS, G. AGGELIS, V. TSAGOU & S. PAVLOU (2005) Dynamics of a two-prey–one-predator system with predator switching regulated by a catabolic repression control-like mode. Ecol Model, 186, 345-357.

2. V. TSAGOU & G. AGGELIS (2006) Growth dynamics of Azospirillum lipoferum at steady and transitory states in the presence of NH4. J Appl Microbiol, 100, 286-295.

3. S. PAPANIKOLAOU, M. GALIOTOU-PANAYOTOU, I. CHEVALOT, M. KOMAITIS, I. MARC & G. AGGELIS (2006) Influence of Glucose and Saturated Free-Fatty Acid Mixtures on Citric Acid and Lipid Production by Yarrowia lipolytica. Current Microbiol, 52, 134-142.

4. S. FAKAS, S. PAPANIKOLAOU, M. GALIOTOU-PANAYOTOU, M. KOMAITIS & G. AGGELIS (2006) Lipids of Cunninghamella echinulata with emphasis to γ-linolenic acid distribution among lipid classes. Appl Microbiol Biotechnol, 73, 676-683.

5. A. KAVADIA, D.V. VAYENAS, S. PAVLOU & G. AGGELIS (2007) Dynamics of free-living nitrogen-fixing bacterial populations in antagonistic conditions. Ecol Mod, 200, 243-253.

6. S. FAKAS, M. GALIOTOU-PANAYOTOU, S. PAPANIKOLAOU, M. KOMAITIS & G. AGGELIS (2007) Compositional shifts in lipid fractions during lipid turnover in Cunninghamella echinulata. Enz Microb Technol, 40, 1321-1327.

7. A. AKTYPIS, M. TYCHOWSKI, G. KALANTZOPOULOS & G. AGGELIS (2007) Studies on bacteriocin (thermophilin T) production by Streptococcus thermophilus ACA-DC 0040 in batch and fed-batch fermentation modes. Antonie Leeuwenhoek-J. of Microbiol, 92, 207-220.

8. S. PAPANIKOLAOU, I. CHEVALOT, M. GALIOTOU-PANAYOTOU, M. KOMAITIS, I. MARC & G. AGGELIS (2007) Industrial derivative of tallow: a promising renewable substrate for microbial lipid, single-cell protein and lipase production by Yarrowia lipolytica. Electron J Biotechnol, 10(3), DOI: 10.2225/vol10-issue3-fulltext-8.

9. S. PAPANIKOLAOU, M. GALIOTOU-PANAYOTOU, S. FAKAS, M. KOMAITIS & G. AGGELIS (2007) Lipid production by oleaginous Mucorales cultivated on renewable carbon sources. Eur J Lipid Sci Technol, 109, 1060-1070.

10. S. PAPANIKOLAOU, S. FAKAS, M. FICK, I. CHEVALOT, M. GALIOTOU-PANAYOTOU, M. KOMAITIS, I. MARC & G. AGGELIS (2008) Biotechnological valorisation of raw glycerol discharged after bio-diesel (fatty acid methyl esters) manufacturing process: Production of 1,3-propanediol, citric acid and single cell oil. Biomass Bioenerg, 32, 60-71.

11. S. PAPANIKOLAOU, M. GALIOTOU-PANAYOTOU, S. FAKAS, M. KOMAITIS & G. AGGELIS (2008) Citric acid production by Yarrowia lipolytica cultivated on olive-mill wastewater-based media. Biores Technol, 99, 2419-2428.

12. S. FAKAS, M. CERTIK, S. PAPANIKOLAOU, G. AGGELIS, M. KOMAITIS & M. GALIOTOU-PANAYOTOU (2008) γ-Linolenic acid production by Cunninghamella echinulata growing on complex organic nitrogen sources. Biores Technol, 99, 5986-5990.

13. S. FAKAS, S. PAPANIKOLAOU, M. GALIOTOU-PANAYOTOU, M. KOMAITIS & G. AGGELIS (2008) Organic nitrogen of tomato waste hydrolysate enhances glucose uptake and lipid accumulation in Cunninghamella echinulata. J Appl Microbiol, 105, 1062-1070.

14. A. KAVADIA, D.V. VAYENAS, S. PAVLOU & G. AGGELIS (2008) Dynamics of free-living nitrogen-fixing bacterial populations and nitrogen fixation in a two-prey–one-predator system. Ecol Mod, 218, 323-338.

15. S. FAKAS, I. PAPAPOSTOLOU, S. PAPANIKOLAOU, C.D. GEORGIOU & G. AGGELIS (2008) Susceptibility to peroxidation of the major mycelial lipids of Cunninghamella echinulata. Eur J Lipid Sci Technol, 110, 1062-1067.

16. S. FAKAS, S. PAPANIKOLAOU, A. BATSOS, M. GALIOTOU-PANAYOTOU, A. MALLOUCHOS & G. AGGELIS (2009) Evaluating renewable carbon sources as substrates for single cell oil production by Cunninghamella echinulata and Mortierella isabellina. Biomass Bioenerg, 33, 573-580.

17. S. PAPANIKOLAOU, G. AGGELIS (2009) Biotechnological valorization of biodiesel derived glycerol waste through production of single cell oil and citric acid by Yarrowia lipolytica. Lipid Technol, 21, 83-87.

18. S. FAKAS, A. MAKRI, M. MAVROMATI, M. TSELEPI, G. AGGELIS (2009) Fatty acid composition in lipid fractions lengthwise the mycelium of Mortierella isabellina and lipid production by solid state fermentation. Biores Technol, 100, 6118-6120.

19. Ch. N. ECONOMOU, A. MAKRI, G. AGGELIS, S. PAVLOU & D.V. VAYENAS (2010) Semi-solid state fermentation of sweet sorghum for the biotechnological production of single cell oil. Biores Technol, 101, 1385–1388.

20. A. MAKRI, S. FAKAS & G. AGGELIS (2010) Metabolic activities of biotechnological interest in Yarrowia lipolytica grown on glycerol in repeated batch cultures. Biores Technol, 101, 2351–2358.

21. S. PAPANIKOLAOU, P. DIAMANTOPOULOU, A. CHATZIFRAGKOU, A. PHILIPPOUSSIS & G. AGGELIS (2010) Suitability of low-cost sugars as substrates for lipid production by the fungus Thamnidium elegans. Energ Fuels, 24, 4078-4086.

22. S. FAKAS, I. KEFALOGIANNI, A. MAKRI, G. TSOUMPELI, G. ROUNI, CH.  GARDELI, S. PAPANIKOLAOU & G. AGGELIS (2010) Characterization of olive fruit microflora and its effect on olive oil volatile compounds biogenesis. Eur J Lipid Sci Technol, 112, 1024-1032.

23. A. CHTZIFRAGKOU, S. FAKAS, M. GALIOTOU-PANAYOTOU, M. KOMAITIS, G. AGGELIS & S. PAPANIKOLAOU (2010) Commercial sugars as substrates for lipid accumulation by Cunninghamella echinulata and Mortierella isabellina fungi. Eur J Lipid Sci Technol, 112, 1048-1057.

24. S. PAPANIKOLAOU & G. AGGELIS (2010) Yarrowia lipolytica: a model microorganism used for production of tailor-made lipids. Eur J Lipid Sci Technol, 112, 639-654.

25. Ch. N. ECONOMOU, G. AGGELIS, S. PAVLOU & D.V. VAYENAS (2010) Modelling of Single-Cell Oil production under nitrogen limited and substrate inhibition conditions. Biotechnol Biong, 108, 1049-55.

26. A. MAKRI, S. BELLOU, M. BIRKOU, K. PAPATREHAS, N.P. DOLAPSAKIS, D. BOKAS, S. PAPANIKOLAOU & G. AGGELIS (2011) Lipids synthesized by micro-algae grown in laboratory and industrial scale bioreactors. Eng Life Sci, 11, 52-58. (Erratum Eng. Life Sci. 2011, DOI: 10.1002/elsc.201190032)

27. A. CHATZIFRAGKOU, A. MAKRI, A. BELKA, S. BELLOU, M. MAVROU, M. MASTORIDOU, P. MYSTRIOTI, G. ONJARO, G. AGGELIS, S. PAPANIKOLAOU (2011) Biotechnological conversions of biodiesel derived waste glycerol by yeast and fungal species. Energy J, 36, 1097-1108.

28. S. PAPANIKOLAOU, A. DIMOU, S. FAKAS, P. DIAMANTOPOULOU, A. PHILIPPOUSSIS, M. GALIOTOU-PANAYOTOU & G. AGGELIS (2011) Biotechnological conversion of waste cooking olive oil into lipid-rich biomass using Aspergillus and Penicillium strains. J Appl Mirobiol, 110, 1138–1150.

29. Ch. N. ECONOMOU, I.A. VASILIADOU, G. AGGELIS, S. PAVLOU & D.V. VAYENAS (2011) Modeling of oleaginous fungal biofilm developed on semi-solid media. Biores Technol, 102, 9697-9704.

30. Ch. N. ECONOMOU, G. AGGELIS, S. PAVLOU & D.V. VAYENAS (2011) Single cell oil production from rice hulls hydrolysate. Biores Technol, 102, 9737-9742.

31. A. CHATZIFRAGKOU, G. AGGELIS, M. KOMAITIS, A.P. ZENG & S. PAPANIKOLAOU (2011) Impact of anaerobiosis strategy and bioreactor geometry on the biochemical response of Clostridium butyricum VPI 1718 during 1,3-propanediol fermentation. Biores Technol, 102, 10625-10632.

32. S. PAPANIKOLAOU & G. AGGELIS (2011) Lipids of oleaginous yeasts. Part I. Biochemistry related with single cell oil production. Eur J Lipid Sci Technol, 113, 1031-1051.

33. S. PAPANIKOLAOU & G. AGGELIS (2011) Lipids of oleaginous yeasts. Part II. Technology and potential applications. Eur J Lipid Sci Technol, 113, 1052-1073.

34. A. KAVADIA, D.V. VAYENAS, S. PAVLOU & G. AGGELIS (2011) Dynamics of a free-living nitrogen-fixing bacteria population lacking of competitive advantage towards an antagonistic population. Open Environmental Engineering J, 4, 190-198.

35. S. BELLOU, A. MOUSTOGIANNI, A. MAKRI & G. AGGELIS (2012) Lipids containing polyunsaturated fatty acids synthesized by Zygomycetes grown on glycerol. Appl Biochem Biotechnol 166, 146–158.

36. A. CHATZIFRAGKOU, G. AGGELIS, CH. GARDELI, M. GALIOTOU-PANAYOTOU, M. KOMAITIS & S. PAPANIKOLAOU (2012) Adaptation dynamics of Clostridium butyricum in high 1,3-propanediol content media. Appl Microbiol Biotechnol, 95, 1541-1552.

37. P. DIAMANTOPOULOU, S. PAPANIKOLAOU, M. KAPOTI, M. KOMAITIS, G. AGGELIS & A. PHILIPPOUSSIS (2012) Mushroom polysaccharides and lipids synthesized in liquid agitated and static cultures. Part I: Screening various mushroom species. Appl Biochem Biotechnol, 167, 536-551.

38. P. DIAMANTOPOULOU, S. PAPANIKOLAOU, E. KATSAROU, M. KOMAITIS, G. AGGELIS & A. PHILIPPOUSSIS (2012) Mushroom polysaccharides and lipids synthesized in liquid agitated and static cultures. Part II: Study of Volvariella volvacea. Appl Biochem Biotechnol, 167, 1890-1906.

39. M. BIRKOU, D. BOKAS & G. AGGELIS (2012) Improving fatty acid composition of lipids synthesized by Brachionus plicatilis in large scale experiments. J. Am. Oil Chem. Soc., 89, 2047-2055.

40. S. BELLOU & G. AGGELIS (2012) Biochemical activities in Chlorella sp. and Nannochloropsis salina during lipid and sugar synthesis in a lab-scale open pond simulating reactor. J Biotechnol, 164, 318-329.

41. S. PAPNIKOLAOU, A. BEOPOULOS, A. KOLETTI, F. THEVENIEAU, A.A. KOUTINAS, J.-M. NICAUD & G. AGGELIS (2013) Importance of the methyl - citrate cycle on glycerol metabolism in the yeast Yarrowia lipolyrica. J Biotechnol, 168, 303-314

42. S. BELLOU, A. MAKRI, D. SARRIS, K. MICHOS, P. RENTOUMI A. CELIK, S. PAPANIKOLAOU & G. AGGELIS (2014) The olive mill wastewater as substrate for single cell oil production by Zygomycetes. J Biotechnol, 170, 50-59.

43. D. SARRIS, L. MATSAKAS, G. AGGELIS, A.A. KOUTINAS, S. PAPANIKOLAOU (2014) Aerated vs non-aerated conversions of molasses and olive mill wastewaters blends into bioethanol by Saccharomyces cerevisiae under non-aseptic conditions. Ind. Crops and Products, 56, 83-93

44. S. BELLOU, A. MAKRI, I.E. TRIANTAPHYLLIDOU, S. PAPANIKOLAOU & G. AGGELIS (2014) Morphological and metabolic shifts of Yarrowia lipolytica induced by alteration of the dissolved oxygen concentration in the growth enviroment. Microbiology, 160, 807-817.

45. A. KOUVELAS, G. AGGELIS, A.A. ALEXOPOULOS & K.C. ANGELOPOULOS (2014) Nitrogen dynamics during growth of sweet sorghum [Sorghum bicolor (L.) Moench] in response to conventional and organic soil fertility management. Australian Journal of Corp Science, 8, 730-737.

46. P. DIAMANTOPOULOU, S. PAPANIKOLAOU, M. KOMAITIS, G. AGGELIS & A. PHILIPPOUSIS (2014) Patterns of major metabolites biosynthesis by different mushroom fungi grown on glucose-based submerged cultures. Bioprocess and Biosystems Engineering, 37, 1385-1400.

47. A. MOUSTOGIANNI, S. BELLOU, I.-E. TRIANTAPHYLLIDOU & G. AGGELIS (2014) Alterations in fatty acid composition of Cunninghamella echinulata lipids induced by orange essential oil. Enviromental Biotechnology, 10, 1-7.

48. S. BELLOU, M.N. BAESHEN, A.M. ELAZZAZY, D. AGGELI, F. SAYEGH & G. AGGELIS (2014) Microalgal lipids biochemistry and biotechnological perspectives. Biotechnology Advances, 32, 1476-1493.

49. A. MOUSTOGIANNI, S. BELLOU, I.-E. TRIANTAPHYLLIDOU & G. AGGELIS (2015) Feasibility of raw glycerol conversion into Single Cell Oil by Zygomycetes under non-aseptic conditions. Biotechnol Bioeng, 112, 827-831. 

50. S.S. TCHAKOUTEU, A. CHATZIFRAGKOU, O. KALANTZI, A.A. KOUTINAS, G. AGGELIS & S. PAPANIKOLAOU (2015) Oleaginous yeast Cryptococcus curvatus exhibits interplay between biosynthesis of intracellular sugars and lipids. European Journal of Lipid Science Technology, 117, 657–672.

51. S.S. TCHAKOUTEU, O. KALANTZI, CH. GARDELI, A.A. KOUTINAS, G. AGGELIS & S. PAPANIKOLAOU (2015) Lipid production by yeasts growing on biodiesel-derived crude glycerol: strain selection and impact of substrate concentration on the fermentation efficiency. Journal of Applied Microbiology, 118, 911-927.

52. Ch. N. ECONOMOU, N. MARINAKIS, M. MOUSTAKA-GOUNI, G. KEHAYIAS, G. AGGELIS & D.V. VAYENAS (2015) Lipid production by the filamentous cyanobacterium Limnothrix sp. growing on a synthetic wastewater in suspended- and attached-growth photobioreactor systems. Annals of Microbiology, 65, 1941–1948.

53. R. ALAKHRAS, S. BELLOU, G. FOTAKI, G. STEPANOU, N.A. DEMOPOULOS, S. PAPANIKOLAOU & G. AGGELIS (2015) Fatty acid lithium salts from Cunninghamella echinulata have cytotoxic and genotoxic effects on HL-60 human leukemia cells. Engineering in Life Sciences, 15, 243–253.

54. E.K.F. ELBESHEHY, A.M. ELAZZAZY & G. AGGELIS (2015) Silver nanoparticles synthesis mediated by newly isolates of Bacillus spp., nanoparticles characterization and their activity against Bean Yellow Mosaic Virus and human pathogens. Frontiers in Microbiology (section Antimicrobials, Resistance and Chemotherapy), 6, 453.

55. F. AROUS, I-E. TRIANTAPHYLLIDOU, T. MECHICHI, S. AZABOU, M. NASRI & G. AGGELIS (2015) Lipid accumulation in the new oleaginous yeast Debaryomyces etchellsii correlates with ascosporogenesis. Biomass and Bioenergy, 80, 307-315.

56. P. DIAMANTOPOULOU, S. PAPANIKOLAOU, G. AGGELIS & A. PHILIPPOUSSIS (2016) Adaptation of Volvariella volvacea metabolism in high carbon to nitrogen ratio media. Food Chem, 196, 272-280.

57. O.N. TSOLCHA, A.G. TEKERLEKOPOULOU, CH.S. AKRATOS, S. BELLOU, G. AGGELIS, M. KATSIAPI, M. MOUSTAKA-GOUNI & D.V. VAYENAS (2016) Treatment of second cheese whey effluents using a Choricystis-based system with simultaneous lipid production. J Chem Technol Biotechnol, 91, 2349–2359.

58. F. SAYEGH, A. ELAZZAZY, S. BELLOU, A. MOUSTOGIANNI, A.I. ELKADY, M.N. BAESHEN & G. AGGELIS (2016) Production of polyunsaturated single cell oils possessing antimicrobial and anticancer properties. Ann Microbiol, 66, 937-948.

59. S. BELLOU, I-E. TRIANTAPHYLLIDOU, D. AGGELI, A-M. MOHAMMED, M-N. BAESHEN & G. AGGELIS (2016) Microbial oils as food additives: recent approaches for improving microbial oil production and its polyunsaturated fatty acid content. Current Opinion in Biotechnology, 37, 24-35.

60. F. ΑROUS, T. MECHICHI, M. NASRI & G. AGGELIS (2016) Fatty acid biosynthesis during the life cycle of Debaryomyces etchellsii. Microbiology – SGM, 162, 1080-1090.

61. F. ΑROUS, F. FΡΙΚΗΑ, I-E. TRIANTAPHYLLIDOU, G. AGGELIS, M. NASRI & T. MECHICHI (2016) Potential utilization of agro-industrial wastewaters for lipid production by the oleaginous yeast Debaryomyces etchellsii. J Clean Prod, 133, 899-909.

62. S. BELLOU, I.-E.TRIANTAPHYLLIDOU, P. MIZERAKIS & G. AGGELIS (2016) High lipid accumulation in Yarrowia lipolytica cultivated under double limitation of nitrogen and magnesium. J Biotechnol, 234, 116–126.

63. M. DOUROU, A. KANCELISTA, P. JUSZCZYK, D. SARRIS, S. BELLOU, I.-E. TRIANTAPHYLLIDOU, A. RYWINSKA, S. PAPANIKOLAOU & G. AGGELIS (2016) Bioconversion of olive mill wastewater into high-added value products. J Clean Prod, 139, 957-969.

64. S. PAPANIKOLAOU, M. RONTOU, A. BELKA, M. ATHENAKI, CH. GARDELI, A. MALLOUCHOS, O. KALANTZI, A.A. KOUTINAS, I.K. KOOKOS, A-P. ZENG & G. AGGELIS (2017) Conversion of biodiesel-derived glycerol into biotechnological products of industrial significance by yeast and fungal strains. Eng Life Sci, 17, 262–281.

65. S.S. TCHAKOUTEU, N. KOPSAHELIS, A. CHATZIFRAGLOU, O. KALANTZI, N.G. STOFOROS, A.A. KOUTINAS, G. AGGELIS & S. PAPANIKOLAOU (2017) Rhodosporidium toruloides cultivated in NaCl-enriched glucose-based media: Adaptation dynamics and lipid production. Eng Life Sci, 17, 237–248

66. F. AROUS, S. AZABOU, I-E. TRIANTAPHYLLIDOU, G. AGGELIS, A. JAOUANI, M. NASRI & T. MECHICHI (2017) Newly isolated yeasts from Tunisian microhabitats: Lipid accumulation and fatty acid composition. Eng Life Sci, 17, 226–236.

67. M. DOUROU, P. MIZERAKIS, S. PAPANIKOLAOU & G. AGGELIS (2017) Storage lipid and polysaccharide metabolism in Yarrowia lipolytica and Umbelopsis isabellina. Appl Microbiol Biotechnol, 101, 7213–7226.

68. P. MIZERAKIS, P. STATHOPOULOU, G. TSIAMIS, M. N. BAESHEN, J. A. MAHYOUB, A.M. ELAZZAZY, S. BELLOU, E. SAKOULOGEARGA, I-E TRIANTAPHYLLIDOU, T. MAZIOTI, P. KATSORIS & G. AGGELIS (2017) Bacterial diversity of the outflows of a Polichnitos (Lesvos, Greece) hot spring, laboratory studies of a Cyanobacterium sp. strain and potential medical applications. Annals of Microbiology, 67, 643-654.

69. CH. GARDELI, M. ATHENAKI, E. XENOPOULOS, A. MALLOUCHOS, A. KOUTINAS, G. AGGELIS & S. PAPANIKOLAOU (2017) Lipid production and characterization by Mortierella (Umbelopsisisabellina cultivated on lignocellulosic sugars. J Appl Microbiol, 123, 1461-1477.

70. M. DOUROU, D. AGGELI, S. PAPANIKOLAOU & G. AGGELIS (2018) Critical steps in carbon metabolism affecting lipid accumulation and their regulation in oleaginous microorganisms. Appl Microbiol Biotechnol, 102, 2509–2523.

71. R. MALIBARI, F. SAYEGH, A.M. ELAZZAZY, M.N. BAESHEN, M. DOUROU & G. AGGELIS (2018) Reuse of shrimp farm wastewater as growth medium for marine microalgae isolated from Red Sea – Jeddah. J Clean Prod, 198, 160-169.

72. I.A. VASILIADOU, S. BELLOU, A. DASKALAKI, L. TOMASZEWSKA-HETMAN, Ch. CHATZIKOTOULA, B. KOMPOTI, S. PAPANIKOLAOU, D.V. VAYENAS, S. PAVLOU & G. AGGELIS (2018) Biomodification of fats and oils and scenarios of adding value on renewable fatty materials through microbial fermentations: Modelling and trials with Yarrowia lipolytica. J Clean Prod, 200, 1111-1129.

73. M. DOUROU, O.N. TSOLCHA, A.G. TEKERLEKOPOULOU, D. BOKAS & G. AGGELIS (2018) Fish farm effluents are suitable growth media for Nannochloropsis gaditana, a polyunsaturated fatty acid producing microalga. Eng Life Sci, 18, 851-860.

74.  A. DASKALAKI, I.A. VASILIADOU, S. BELLOU, L. TOMASZEWSKA-HETMAN, Ch. CHATZIKOTOULA, B. KOMPOTI, S. PAPANIKOLAOU, D.V. VAYENAS, S. PAVLOU & G. AGGELIS (2018) Data on cellular lipids of Yarrowia lipolytica grown on fatty substrates. J Clean Prod, 21, 1037-1044.

75. A. DASKALAKI, N. PERDIKOULI, D. AGGELI & G. AGGELIS (2019) Laboratory evolution strategies for improving lipid accumulation in Yarrowia lipolytica.  Appl Microbiol Biotechnol, 10.1007/s00253-019-10088-7.

76. Μ. KOTHRI, M. MAVROMATI, A M ELAZAZY, M N BAESHEN, T AA MOUSSA & G AGGELIS (2019) Microbial sources of polyunsaturated fatty acids (PUFAs) and the prospect of organic residues and wastes as growth media for PUFA-producing microorganisms. FEMS Microbiology Letters, fnaa028.

77. M. LLAMAS, M. DOUROU, C. GONZALEZ-FERNANDEZ, G. AGGELIS & E. TOMAS-PEJO (2020) Screening of oleaginous yeasts for lipid production using volatile fatty acids as substrate. Biomass Bioenergy, 138, 105553.

78. M. DOUROU, P. DRITSAS, M.N. BAESHEN, A.M. ELAZZAZY, A. ALFERGAH & G. AGGELIS (2020) High-added value products from microalgae and prospects of aquaculture wastewaters as microalgae growth media. FEMS Microbiology Letters, fnaa081.

79. V. PATRINOU, O. TSOLCHA, T. TATOULIS, N. STEFANIDOU, M. DOUROU, M. MOUSTAKA-GOUNI, G. AGGELIS & A. TEKERLEKOPOULOU (2020) Biotreatment of poultry waste coupled with biodiesel production using suspended and attached growth microalgal-based systems. Sustainability, 12, 5024.

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